appler
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Appler has an issue with 486s
In 86Box, an emulated machine set up as a 486 is unable to use Appler (hard freezes). I reported it as a bug and they said it works on an emulated Pentium.
Originally Appler worked on 8088/CGA. Later it required EGA. Later still Ivo & I wrote code to accumulate 6502 clock cycles in the high order 16 bits of eax, which of course requires a 32-bit CPU. The accumulated value is used to sync up with real time whenever the 6502 program touches I/O memory, e.g. $C030 (for sound).
If you search in the source code, you'll find (very few) other places where 32-bit registers are used.
I don't recall making any changes that require Pentium, but it's possible that I have.