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Font Headings is set to "Source Serif" and not "Source Serif 4"
The fonts linked in the README.md are Source Serif 4 and DM Sans. While DM Sans is correct in theme.css, Source Serif 4 is improperly listed, leading to headings not using the correct font.
To fix, either replace Source Serif with Source Serif 4, or add Source Serif 4 as an additional font.