web-ext-translator
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regional variant separator
https://developer.mozilla.org/en-US/docs/Mozilla/Add-ons/WebExtensions/Internationalization#Providing_localized_strings_in__locales
However, in the directories under _locales, the separator must be an underscore: "en_US".
Firefox accepts both - and _, but Chrome only accepts _.
The desktop version of this app saves folders with an underscore and if you export the files as ZIP, the folders use underscores as well. it's only in the editor that the standard - is used.
I guess I could improve the prepared text for github issues to include the directory name, so that add-on authors don't make this mistake (I stumbled upon this myself just yesterday).